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Smithsonian National Museum of Natural History
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Department of Paleobiology

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  • Robert J. Emry
  • Curator of Cenozoic Mammals, emeritus
  • Phone:   202-633-1323
  • Fax:   202-786-2832
  • E-mail Address:   emryr
  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 121
    Washington, DC 20013-7012
  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th & Constitution NW
    Washington, DC 20560-0121
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Education

Ph.D. Columbia University
B.S. Colorado State University

Research Interests

Evolution, Relationships, Biochronology and Paleogeography of Cenozoic Mammals.
Broadly, my research is centered on fossil mammals, more specifically the land mammals of the late Eocene and Oligocene epochs (about 45 to 30 million years ago) in the northern hemisphere. I'm interested in understanding how the composition of whole land mammal faunas changes through time, both by evolutionary processes and by recombining through geographic dispersal and interchange. The late Eocene to Oligocene interval was a time of relatively rapid cooling on a global scale (palm trees and crocodiles at 80 degrees north latitude in the Eocene to ice caps at the poles in the Oligocene). This shift to more temperate, and more seasonal, climates on land appears to coincide with changes in oceanic circulation patterns caused by changes in positions of the continents and the connections between them. This same interval was also a time of increased interchange of land mammals between continents, especially between Eurasia and North America. For much of the last decade I've collected and studied mammalian faunas representing this time interval in China and Central Asia, improving identifications of the elements of these faunas and the biochronology based on them, and comparing and correlating with coeval North American land mammal faunas that I'm more familiar with through earlier work. Better time constraints on these episodes of intercontinental interchange of mammals will permit us to see how these episodes correlate with, and are affected by, other major changes in Earth history.

Research Assistants, Current and Former Post-doctoral Fellows, and Students

Post-Doctoral:

  • Kenneth D. Rose (1980-1981), Currently Professor of Anatomy, Department of Cell Biology and Anatomy, Johns Hopkins University, Baltimore.
  • Timothy Heaton (1988-1989), Currently Professor of Geology,
    Department of Earth Sciences, University of South Dakota, Vermillion.
  • Wang Ban-Yue (1989-1990), Currently Senior Researcher, Institute for Vertebrate Paleontology and Paleoanthropology, Beijing.
  • Jin Meng (1991-1992), Currently Curator of Fossil Mammals,
    Department of Vertebrate Paleontology, American Museum of Natural History, New York.

Pre-Doctoral

  • Luke Holbrook (1994), Currently Associate Professor, Department of Biological Sciences Rowan University, Glassboro, NJ

Summer Interns

  • Chad Schennum (2000)

Affiliations

  • Research Associate, Department of Vertebrate Paleontology, American Museum of Natural History (1971-present).

Editorial Service

  • Co-Editor, Journal of Vertebrate Paleontology, 1990-1993.
  • Editor for Special Publications, Society of Vertebrate Paleontology, 1993-present.
  • Regional (Eastern) Editor, Society of Vertebrate Paleontology News Bulletin, 1971-1976.

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Publications

publicationsLucas, Spencer G., Holbrook, Luke T., and Emry, Robert J. 2003. Isectolophus (Mammalia, Perissodactyla) from the Eocene of the Zaysan Basin, Kazakstan, and its biochronological significance. Journal of Vertebrate Paleontology 23 (1):238-243.

Asher, Robert J., McKenna, Malcolm C., Emry, Robert J., Tabrum, Alan R., and Kron, Donald G. 2002. Morphology and Relationships of Apternodus and other Extinct Zalambdodont, Placental Mammals. Bulletin of the American Museum of Natural History, No. 73:117 pp., 60 figs., 7 tables.

Bayshashov, B. U., Lucas, S. G., and Emry, R. J. 2002. Novye danniye k paleontologicheskim issledovaniyat v mestonakhozhdeniyakh Aktau (Dzungarskii Alatau) [New data on the paleontological investigations of the Aktau Locality (Dzhungarian Alatau)]; pages 14-15 in Zoologicheskiye Issledovaniya v Kazakhstanye: Sovremennoye Sostoyaniye i Perspektivyi, Materialy Mezhdunarodnoi Nauchnoi Konferentsii, 19-21 Marta 2002 g. Almaty [Zoological Investigations in Kazakhstan: State of Art and Perspectives. Materials of the International Scientific Conference, 19021 March, 2002, Almaty]. [abstract]

Bayshashov, Bolat, Tyutkova, Lyubov A., Lucas, Spencer G., and Emry, Robert J. 2002. New Mammal Finds and the Early Miocene Age of the Askazansor Locality, Kazakstan. Journal of Vertebrate Paleontology, Vol. 22 (supplement to no. 3):34A

Emry, Robert J. (editor). 2002. Cenozoic Mammals of Land and Sea: Tributes to the Career of Clayton E. Ray. Smithsonian Contributions to Paleobiology, no. 93, vi + 372 pages, 197 figures, 11 plates, 46 tables.

Emry, Robert J. 2002. Preface; page v in, Robert J. Emry (Editor), Cenozoic Mammals of Land and Sea: Tributes to the Career of Clayton E. Ray. Smithsonian Contributions to Paleobiology, no. 93.

Emry, Robert J., and Lucas, Spencer G. 2002. Brontothere (Mammalia, Perissodactyla) Footprints From the Eocene of the Ily Basin, Kazakstan. Journal of Vertebrate Paleontology, Vol. 22 (supplement to no. 3):51A.

Erbayeva, M. A., Emry, R. J., and Tyutkova, L. A. 2002. LAGOMORPHS pages 20-22 in Zoologicheskiye Issledovaniya v Kazakhstanye: Sovremennoye Sostoyaniye i Perspektivyi, Materialy Mezhdunarodnoi Nauchnoi Konferentsii, 19-21 Marta 2002 g. Almaty [Zoological Investigations in Kazakhstan: State of Art and Perspectives. Materials of the International Scientific Conference, 19021 March, 2002, Almaty]. [abstract]

Eshelman, Ralph E., Emry, Robert J., Domning, Daryl P., and Bohaska, David J. 2002. Biography and Bibliography of Clayton Edward Ray; pages 1-13 in, Robert J. Emry, (Editor), Cenozoic Mammals of Land and Sea: Tributes to the Career of Clayton E. Ray. Smithsonian Contributions to Paleobiology, no. 93.

Gaudin, Timothy A., and Emry, Robert J. 2002. The Late Eocene Pangolin Patriomanis from North America, and a New Genus of Pangolin from the Late Eocene of Nei Mongol, China (Mammalia, Pholidota). Journal of Vertebrate Paleontology, Vol. 22 (supplement to no. 3):57A

Grady, F., Arroyo-Cabrales, J., and Garton, E. R. 2002. The Northernmost Occurrence of the Pleistocene Vampire Bat Desmodus stocki (Chiroptera: Phyllostomatidae: Desmodontinae) in Eastern North America; pages 73-75 in, Robert J. Emry, (Editor), Cenozoic Mammals of Land and Sea: Tributes to the Career of Clayton E. Ray. Smithsonian Contributions to Paleobiology, no. 93.

McKenna, Malcolm C., Asher, Robert, Emry, Robert, Tabrum, Alan, and Kron, Donald. 2002. The Search for Extinct Relatives of Modern Mammals: The Case of Soricids and Apternodus. Journal of Vertebrate Paleontology, Vol. 22 (supplement to no. 3):86A.

Miyata, Kazunori, and Emry, Robert. 2002. New Information on the Dental Morphology of Trogosus Material from the Bridger Formation, Uinta County, Wyoming, and its Implication to the Evolutionary History of Tillodonts. Journal of Vertebrate Paleontology, Vol. 22 (supplement to no. 3):89A

Storch, Gerhard D., Emry, R. J., Gaudin, T. J., and Rose, K. D. 2002. Origin and Relationships of Xenarthra and Pholidota. Journal of Vertebrate Paleontology, Vol. 22 (supplement to no. 3):111A.

Tleuberdina, P. A., Emry, R. J., and Semenov, Y. A. 2002. CARNIVORES, pages 33-34 in Zoologicheskiye Issledovaniya v Kazakhstanye: Sovremennoye Sostoyaniye i Perspektivyi, Materialy Mezhdunarodnoi Nauchnoi Konferentsii, 19-21 Marta 2002 g. Almaty [Zoological Investigations in Kazakhstan: State of Art and Perspectives. Materials of the International Scientific Conference, 19021 March, 2002, Almaty]. [abstract]

Tyutkova, L. A., Emry, R. J., and Lucas, S. G. 2002. Drevniye khomyakobrazniye Zaysana [Ancient hamsters of Zaysan]; pages 39-40 in Zoologicheskiye Issledovaniya v Kazakhstanye: Sovremennoye Sostoyaniye i Perspektivyi, Materialy Mezhdunarodnoi Nauchnoi Konferentsii, 19-21 Marta 2002 g. Almaty [Zoological Investigations in Kazakhstan: State of Art and Perspectives. Materials of the International Scientific Conference, 19021 March, 2002, Almaty]. [abstract]

Tyutkova, L. A., Erbayeva, M. A., Emry, R. J., and Lucas, S. G. 2002. Miotsenovaya mikroteriofauna Zaysana I yeyo biostratigraficheskoye znacheniye [Miocene microtheriofauna of Zaysan and its biostratigraphic significance]; pages 37-38 in Zoologicheskiye Issledovaniya v Kazakhstanye: Sovremennoye Sostoyaniye i Perspektivyi, Materialy Mezhdunarodnoi Nauchnoi Konferentsii, 19-21 Marta 2002 g. Almaty [Zoological Investigations in Kazakhstan: State of Art and Perspectives. Materials of the International Scientific Conference, 19021 March, 2002, Almaty]. [abstract]

Emry, Robert J. and William W. Korth. 2001. Douglassciurus, new name for Douglassia Emry and Korth, 1996, not Douglassia Bsrtsch, 1934. Journal of Vertebrate Paleontology, vol. 21(2), page 400.
Spencer G. Lucas and Robert J. Emry. 2001. Sharamynodon (Mammalia: Perissodactyls) from the Eocene of the Ily Basin, Kazakstan and the Antiquity of Asian Amynodonts. Proceedings of the Biological Society of Washington, vol. 114(2), pages 517-525.

Lucas, Spencer G., Emry, R.J., Chkhikvadze, V.,Bayshashov, B., Tjutkova,L.A., Tleuberdina, P.A., and Zhamangara, A. 2000. Upper Cretaceous-Cenozoic lacustrine deposits of the Zaysan Basin, eastern Kazakstan. IN: E. H. Gierlowski-Kordesch and K. R. Kelts (Eds.), Lake Basins Through Space and Time: AAPG Studies in Geology 46: 335-340.

Lucas, Spencer G., Robert J. Emry, Lyubov A. Tyutkova and Bolat B. Bayshashov. Fossil Mammals and the Eocene-Oligocene Boundary in the Zaysan Basin, Kazakstan. Selevinia (The Zoological Journal of Kazakstan) 1996-1997 (but published in 1999), pages 64-70. [in Russian, with English summary]

Emry, R. J., Lucas, S. G., Tyutkova, L., and Wang, B. "The Ergilian-Shandgolian (Eocene-Oligocene) transition in the Zaysan Basin, Kazakhstan." In Beard, K. C. and Dawsons, M. R. (eds.), Dawn of the Age of Mammals in Asia. Bulletin of Carnegie Museum of Natural History, 34. pp. 298-312.

Emry, R. J., Tyutkova, L., Lucas, S. G., and Wang, B. "Rodents of the Middle eocene Chinzhaly Fauna of Eastern Kazakhstan." Journal of Vertebrate Paleontology 18(1), pp. 218-227.

Emry, R. J. and Eshelman, R. A. "The Early Hemingfordian (Early Miocene) Pollack Farm Local Fauna: First Tertiary Land Mammals Described from Delaware." In Benson, R. N., ed., Geology and paleontology of the lower Miocene Pollack Farm Fossil Site, Delaware: Delaware Geological Survey Special Publication No. 21, p. 153-173.

Lavarov, A. V. and Emry, R. J. "Hyaenodon venturae (Hyaenodontidae, Creodonta, Mammalia) from the Early Chadronian (Latest Eocene) of Wyoming." Journal of Paleontology 72(4), pp. 752-757.
Lucas, S. G., Emry, R. J., and Foss, S. E. "Taxonomy and Distribution of Daeodon, an Oligocene-Miocene Entelodont (Mammalia: Artiodactyla) from North America." Proceedings of the Biological Society of Washington, 111(2), pp. 425-435.

Lucas, S. G. , Kordikova, E. G. and Emry, R. J. " Oligocene stratigraphy, sequence stratigraphy and mammalian biochronology north of the Aral Sea, central Kazakhstan." In Beard, K. C. and Dawsons, M. R. (eds.), Dawn of the Age of Mammals in Asia. Bulletin of Carnegie Museum of Natural History, 34. pp. 313-348.

Domning, D. P., Emry, R. J., Portell, R. W., Donovan, S. K., and Schindler, K. S. "Oldest West Indian land mammal: rhinocerotoid ungulate from the Eocene of Jamaica." Journal of Vertebrate Paleontology 17(4):638-641.

Emry, R. J., Wang, B., Tjutkova, L.A., and Lucas, S. G. "A late Eocene eomyid rodent from the Zaysan Basin of Kazakhstan." Journal of Vertebrate Paleontology 17:229-234.

Korth, W. W. and Emry, R. J. "The skull of Anchitheriomys and a new subfamily of beavers (Castoridae, Rodentia)." Journal of Paleontology 71:343-347.

Lucas, S. G., Emry, R. J., and Bayshashov, B. U. "Eocene Perissodactyla from the Shinzhaly River, easteran Kazakhstan." Journal of Vertebrate Paleontology 17:235-246.

Emry, R. J., and Korth, W. W. "The Chadronian Squirrel "Sciurus" jeffersoni Douglas, 1901: a New Generic Name, New Material, and its Bearing on the Early Evolution of Sciuridae (Rodentia)." Journal of Vertebrate Paleontology 16 (4) (1996): 770-775.

Emry, R. J. "Cylindrodontidae." In Prothero, D. R., and Emry, R. J., editors. The Terrestrial Eocene-Oligocene Transition in North America (1996): 399-416. Cambridge: Cambridge University Press, 688 pages.

Heaton, T. H., and Emry, R. J. "Leptomerycidae." In Prothero, D. R., and Emry, R. J., editors. The Terrestrial Eocene-Oligocene Transition in North America (1996): 581-608. Cambridge: Cambridge University Press, 688 pages.

Lucas, S. G., and Emry, R. J. "Early Record of Indricothere (Mammalia: Perissodactyla: Hyracondontidae) from the Aral Sea Region of Western Kazakhstan." Proceedings of the Biological Society of Washington 109 (2)(1996): 391-396.

Lucas, S. G., and Emry, R. J. "Late Eocene Entelodonts (Mammalia: Artiodactyla) from Inner Mongolia, China." Proceedings of the Biological Society of Washington 109 (2)(1996): 397-405.

Lucas, S. G., and Emry, R. J. "Biochronological Significance of Amynodontidae (Mammalia, Perissodactyla) from the Paleogene of Kazakhstan." Journal of Paleontology 760 (4)(1996): 691-696.

Lucas, S. G., Emry, R. J., and Purdy, R. J. "Marine Fossil Shark (Chondrichthyes) from Nonmarine Eocene Sediments, Northeastern Kazakhstan." Proceedings of the Biological Society of Washington 109 (2)(1996): 349-352.

Lucas, S. G., Emry, R. J., and Bayshashov, B. U. "Zaisanamydon, a late Eocene amynodontid (Mammali, Perissodactyla) from Kazakhstan and China." Tertiary Research 17(1+2):51-58.

Prothero, D. R., and Emry, R. J., editors. The Terrestrial Eocene-Oligocene Transition in North America (1996). Cambridge: Cambridge University Press, 688 pages.

Prothero, D. R., and Emry, R. J. "Summary." In Prothero, D. R., and Emry, R. J., editors. The Terrestrial Eocene-Oligocene Transition in North America (1996): 664-683. Cambridge: Cambridge University Press, 688 pages.

Emry, R. J. "Mammalian Range Zones in the Chadronian White River Formation at Flagstaff Rim, Wyoming." In Eocene-Oligocene Climatic and Biotic Evolution (1992): 106-115. Edited by D. R. Prothero and W. A. Berggren. Princeton, New Jersey: Princeton University Press, 588 pages.

Emry, R. J., and Korth, W. W. "Evolution in Yoderimyinae (Eomyidae: Rodentia), with New Material from the White River Formation (Chadronian) at Flagstaff Rim, Wyoming." Journal of Paleontology 67 (6)(1993): 1047-1057.

Rose, K. D., and Emry, R. J. "Relationships of Xenarthra, Pholidota, and Fossil `Edentates': the Morphological Evidence." In Mammal Phylogeny; Placentals (1993): 81-102. Edited by F. S. Szalay; M. C. McKenna; and M. J. Novacek. New York: Springer-Verlag New York, Inc., 321 pages.

Emry, R. J. "Introduction - A Biographic Sketch of Morris F. Skinner." In The Species, Genera, and Tribes of the Living and Extinct Horses of the World, 1758-1966; from the Work of Morris F. Skinner. Edited by J. R. Mcdonald, M. L. Macdonald, and L. M. Toohey. Dakoterra 4 (1992): v-viii. Rapid City, South Dakota: South Dakota School of Mines and Technology, 429 pages.

Rose, K. D., and Emry, R. J. "Relationships of Xenarthra, Pholidota, and Fossil "Edentates:" the Morphological Evidence." In Mammal Phylogeny; Placentals (1993): 81-102. Edited by F. S. Szalay, M. Novacek, and M. C. McKenna. New York: Springer-Verlag, 336 pages.

Rose, K. D., Emry, R. J., and Gingerich, P. D. "Skeleton of Alocodontulum atopum, an Early Eocene Epoicotheriid (Mammalia, Palaeanodonta) from the Bighorn Basin, Wyoming." Contributions from the Museum of Paleontology, The University of Michigan 28 (10)(1992): 221-245.

Emry, R. J. "Mammalian Range Zones in the Chadronian White River Formation at Flagstaff Rim, Wyoming." In Eocene-Oligocene Climatic and Biotic Evolution (1992): 106-115. Edited by D. R. Prothero and W. A. Berggren. Princeton, New Jersey: Princeton, University Press, 568 pages.

Korth, W. W., and Emry, R. J. "The Skull of Cedromus and a Review of the Cedromurinae (Rodentia: Sciuridae)." Journal of Paleontology 65 (6) (1991): 984-994.

Korth, W. W., Wahler, J J., and Emry, R. J. "A New Species of Heliscomys and Recognition of the Family Heliscomyidae (Geomyoidea: Rodentia)." Journal of Vertebrate Paleontology 11 (2)(1991): 247-256.

Wang, B., and Emry, R. J. "Eomyidae (Rodentia: Mammalia) from the Oligocene of Nei Mongol, China." Journal of Vertebrate Paleontology 11 (3)(1991): 370-377.

Emry, R. J. "A Tiny New Eocene Ceratomorph and Comments on "Tapiroid" Systematics." Journal of Mammalogy 70 (4)(1989): 794-804.

Emry, R. J. "Mammals of the Bridgerian (Middle Eocene) Elderberry Canyon Local Fauna of Eastern Nevada." In Dawn of the Age of Mammals in the Northern Part of the Rocky Mountains Interior, North America: Geological Society of America Special Paper 243 (1990): 187-210. Edited by T. M. Bown and K. D. Rose. Boulder, Colorado: Geological Society of America, 245 pages.

Emry, R. J., and Korth, W. W. "Rodents of the Bridgerian (Middle Eocene) Elderberry Canyon Local Fauna of Eastern Nevada." Smithsonian Contributions to Paleobiology Number 67 (1990). Washington, D. C.: Smithsonian Institution Press, 14 pages.